1982 – PhD – University of Chicago – Evolutionary Biology
1976 – BS – University of California, Berkeley – Zoology
Evolutionary biology, ecology and conservation biology of amphibians and reptiles. Recent research projects include comparative phylogeography of amphibians and reptiles in California and the central U.S., systematics of freshwater turtles and tortoises in Australia, California, and the rest of the globe, and conservation genetics of endangered California amphibians and reptiles. Recently, we have focused a great deal of ecological and genetical work on the California tiger salamander, an endangered species native to central California grassland habitat.